The ovulate cone of the pine can take 3 years to mature. Once a pollen grain from a male cone (microstrobilus) has landed on the female cone (megastrobilus), fertilization of the egg takes a year! After fertilization, the resulting embryo must mature into a seed.
These are studies of immature female cones, specifically the archegonia, where the ovules are found. The ovules appear as eyes in the faces of the manipulated images.